Arachnology @ AMNH

Scientific Publications


Botero-Trujillo, R., Ott, R., Mattoni, C.I., Nime, M.F., and Ojanguren-Affilastro, A.A. 2019. Two new species of the sun-spider genus Gaucha from Argentina and Brazil (Solifugae, Mummuciidae). Zootaxa 4551(2): 180–194.PDF

Botero-Trujillo, R., Lagos-Silnik, S., and Fernandez-Campon, F. 2019. Curanahuel aconcagua, a new genus and species of sun-spider (Solifugae: Mummuciidae) from the Cuyan High Andean biogeographic province of Argentina. The Journal of Arachnology 47(3): 351–359.PDF

Carvalho, L.S., and Botero-Trujillo, R. 2019. On the sun-spiders of the ibirapemussu species-group of the genus Gaucha Mello-Leitão, 1924 (Solifugae, Mummuciidae), with description of a new species. Zootaxa 4700(2): 289–298.PDF

Esposito, L.A. and Prendini, L. 2019. Island ancestors and New World Biogeography: A case study from the Scorpions (Buthidae, Centruroidinae). Scientific Reports 9: 3500. PDF

Monjaraz-Ruedas, R., Francke, O.F., and Prendini, L. 2019. Systematics of the Short-tailed Whipscorpion Genus Stenochrus Chamberlin, 1922 (Schizomida: Hubardiidae) with descriptions of six new genera and five new species. Bulletin of the American Museum of Natural History 435: 1–91. PDF

Ramirez, M.J., Grismado, C.J., Ubick, D., Ovtsharenko, V., Cushing, P.E., Platnick, N., Wheeler, W.C., Prendini, L., Crowley, L. and Horner, N. 2019. Myrmecicultoridae, a New Family of Myrmecophilic Spiders from the Chihuahuan Desert (Araneae: Entelegynae). American Museum Novitates 3930: 1–24. PDF


Barrales Alcalá, D.A., Francke, O.F., and Prendini, L. 2018. Systematic revision of the giant North American vinegaroons, genus Mastigoproctus (Thelyphonida: Thelyphonidae). Bulletin of the American Museum of Natural History 418: 1–62. PDF

Esposito, L.A., Yamaguti, H.Y., Pinto-da-Rocha, R. and Prendini, L. 2018. Plucking with the plectrum: Phylogeny of the New World buthid scorpion subfamily Centruroidinae Kraus, 1955 (Scorpiones: Buthidae) reveals evolution of three pecten-sternite stridulation organs. Arthropod Systematics and Phylogeny 76(1): 87–122. PDF

González-Santillán, E., and Prendini, L. 2018. Systematic Revision of the North American Syntropine Vaejovid Scorpion Genera Balsateres, Kuarapu, and Thorellius, With Descriptions of three New Species. Bulletin of the American Museum of Natural History 420: 1–81. PDF

Loria, S.F. and Prendini, L. 2018. Ultrastructural comparison of the eyespot and ocelli of scorpions, and implications for the systematics of Chaerilidae Pocock, 1893. Zoologischer Anzeiger 273: 183-191. PDF

Miranda, G.S., Giupponi, A.P.L., Prendini, L. and Scharff, N. 2018. Weygoldtia, a new genus of Charinidae Quintero, 1986 (Arachnida, Amblypygi) with a reappraisal of the genera in the family. Zoologischer Anzeiger 273: 23-32. PDF

Prathapan, K.D. et al. [list of authors including Prendini, L. 2018. When the cure kills—CBD limits biodiversity research. Science 360(6396): 1405-1406. PDF

Schaffrath, S., Prendini, L., Reinhard, P. 2018. Intraspecific venom variation in southern African scorpion species of the genera Parabuthus, Uroplectes and Opistophthalmus (Scorpiones: Buthidae, Scorpionidae). Toxicon 144: 83-90. PDF

Valdez-Mondragón, A., Francke, O.F. and Botero-Trujillo, R. 2018. New morphological data for the order Ricinulei with the description of two new species of Pseudocellus (Arachnida: Ricinulei: Ricinoididae) from Mexico. Journal of Arachnology 46: 114-132. PDF


Esposito, L.A, Yamaguti, H.Y., Souza, C.A., Pinto-da-Rocha, R. and Prendini, L. 2017. Systematic revision of the Neotropical club-tailed scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, revalidation of Heteroctenus, and description of two new genera and three new species (Buthidae: Rhopalurusinae) 415: 1–134 Bulletin of the American Museum of Natural History 415: 1–134. PDF

Rogers, D.C., et al. [list of authors including Prendini, L.] 2017. Images are not and should not ever be type specimens: A rebuttal to Garraffoni & Freitas. Zootaxa 4269(4): 455–459. PDF

Rubin, M., Lamsdell, J.C., Prendini, L., and Hopkins, M.J. 2017. Exocuticular hyaline layer of sea scorpions and horseshoe crabs suggests cuticular fluorescence is plesiomorphic in chelicerates Journal of Zoology (London) 303(4): 245–253. PDF


Adams, A., Marais, E., Turner, S., Prendini, L., and Pinshow, B. 2016. Similar burrow architecture of three arid-zone scorpion species implies similar ecological function. Natuurwissenschaften (The Science of Nature) 103: 56. DOI 10.1007/s00114-016-1374-z. PDF

Amorim, D.S., Santos, C.M.D., Krell, F.-T., Dubois, A., Nihei, S.S., Oliveira, O.M.P., Pont, A., Song, H., Verdade, V.K., Fachin, D.A., Klassa, B., Lamas, C.J.E., Oliveira, S.S., De Carvalho, C.J.B., Mello-Patiu, C.A., Hajdu, E., Couri, M.S., Silva, V.C., Capellari, R.S., Falaschi, R.L., Feitosa, R.M., Prendini, L., Pombal Jr., J.P., Fernández, F., Rocha, R.M., Lattke, J.E., Caramaschi, U., Duarte, M., Marques, A.C., Reis, R.E., Kurina, O., Takiya, D.M., Tavares, M., Fernandes, D.S., Franco, F.L., Cuezzo, F., Paulson, D., Guénard, B., Schlick-Steiner, B.C., Arthofer, W., Steiner, F.M., Fisher, B.L., Johnson, R.A., Delsinne, T.D., Donoso, D.A., Mulieri, P.R., Patitucci, L.D., Carpenter, J.M., Herman, L. & Grimaldi, D. 2016. Timeless standards for species delimitation. Zootaxa 4137(1): 121–128. PDF

Ceccarelli, F.S., Ojanguren-Affilastro, A.A., Ramírez, M.J., Ochoa, J.A., Mattoni, C.I., and Prendini, L. 2016. Andean uplift drives diversification of the bothriurid scorpion genus Brachistosternus. Journal of Biogeography. [Early Edition: doi:10.1111/jbi.12760] PDF

Ceriaco, L.M.P., Gutiérrez, E.E., Dubois, A., et al. [493 coauthors including Prendini, L.] 2016. Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa 4196(3): 435–445. PDF

Franz-Guess, S., Klußman-Fricke, B.-J., Wirkner, C.S., Prendini, L., and Starck, J.M. 2016. Morphology of the tracheal system of camel spiders (Chelicerata: Solifugae) based on micro-CT and 3D-reconstruction in exemplar species from three families. Arthropod Structure & Development 45: 440–451. PDF

González-Santillán, E. and Prendini, L. 2016. Systematic revision of the North American syntropine vaejovid scorpion genera Maaykuyak, Syntropis, and Vizcaino, with description of the adults of Syntropis williamsi. Bulletin of the American Museum of Natural History 405: 1–67. PDF

Janion-Scheepers, C., Measey, J., Braschler, B., Chown, S.L., Coetzee, L., Colville, J., Dames, J., Davies, A.B., Davis, A., Davies, S., Dippenaar-Schoeman, A., Duffy, G., Fourie, D., Griffiths, C., Haddad, C., Hamer, M., Herbert, D., Hugo-Coetzee, L.E.A., Jacobs, A., Jansen van Rensburg, C., Lamani, S., Lotz, L.N., Louw, S., Lyle, R., Malan, A., Marais, M., Neethling, J.-A., Nxele, T., Plisko, D., Prendini, L., Swart, A., Theron, P., Truter, M., Ueckermann, E., Uys, V.M., Villet, M.H., Willows-Munrow, S., and Wilson, J.R.U. 2016. Soil biota in a megadiverse country: Current knowledge and future directions in South Africa. Pedobiologia – Journal of Soil Ecology 59: 129–174. PDF

Maquart, P.O., Réveillion, F., Prendini, L., Burger, M., Fisher, B.L., and Van Noort, S. (2016). New distribution records for African whip spiders (Arachnida: Amblypygi). African Entomology 24(1): 245-246. PDF

Ojanguren-Affilastro, A.A., Mattoni, C.I., Ochoa, J.A., Ramírez, M.J., Ceccarelli, F.S., Prendini, L. 2016. Phylogeny, species delimitation and convergence in the South American bothriurid scorpion genus Brachistosternus Pocock 1893: Integrating morphology, nuclear and mitochondrial DNA. Molecular Phylogenetics and Evolution 94: 159–170. PDF

Prendini, L. 2016. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History 407: 1–66. PDF

Wheeler W.C., Coddington, J.A., Crowley, L.M., Dimitrov, D., Goloboff, P.A., Griswold, C.E., Hormiga, G., Prendini, L., Ramírez, M.J., Sierwald, P., Almeida-Silva, L., Alvarez-Padilla, F., Arnedo, M.A., Benavides Silva, L.R., Benjamin, S.P., Bond, J.E., Grismado, C.J., Hasan, E., Hedin, M., Izquierdo, M.A., Labarque, F.M., Ledford, J., Lopardo, L., Maddison, W.P., Miller, J.A., Piacentini, L.N., Platnick, N.I., Polotow, D., Silva-Dávila, D., Scharff, N., Szuts, T., Ubick, D., Vink, C.J., Wood, H.M., and Zhang, J. 2016. The spider tree of life: Phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics DOI: 10.1111/cla.12182. PDF


Amr, Z., Abed, O.A., Al Share, T., Hamidan, N., and Prendini, L. 2015. New records of Jordanian scorpions. Jordan Journal of Natural History 2: 30–38. PDF

Bird, T.L., Wharton, R.A., and Prendini, L. 2015. Cheliceral morphology in Solifugae (Arachnida): Primary homology, terminology, and character survey. Bulletin of the American Museum of Natural History 394: 1–355. PDF

Cushing, P.E., Graham, M.R., Prendini, L., and Brookhart, J.O. 2015. A multilocus molecular phylogeny of the endemic North American camel spider family Eremobatidae (Arachnida: Solifugae). Molecular Phylogenetics and Evolution 92: 280–293. PDF

González-Santillán, E. and Prendini, L. 2015. Phylogeny of the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, based on morphology, mitochondrial and nuclear DNA. Cladistics 31(4): 341–405. PDF

González-Santillán, E. and Prendini, L. 2015. Systematic revision of the North American syntropine vaejovid scorpions with a subaculear tubercle, Konetontli González-Santillán and Prendini, 2013. Bulletin of the American Museum of Natural History 397: 1–78. PDF

Mattoni, C.I., García-Hernández, S., Botero-Trujillo, R., Ochoa, J.A., Ojanguren-Affilastro, A.A., Pinto-da-Rocha, R., and Prendini, L. 2015. Scorpion sheds ‘tail’ to escape. First report of autotomy in scorpions (Buthidae: Ananteris). PLoS One10(1): e0116639 (15 pp.). PDF

Prendini, L. 2015. A remarkably small species of Uroplectes Peters, 1861 (Scorpiones: Buthidae) endemic to the Succulent Karoo of the Northern Cape Province, South Africa. African Invertebrates 56(2): 499–513. PDF

Prendini, L. 2015. Three new Uroplectes (Scorpiones: Buthidae) with punctate metasomal segments from tropical central Africa. American Museum Novitates 3840: 1–32. PDF

Talal, S., Tesler, I., Sivan, J., Ben-Shlomo, R., Tahir, H.M., Prendini, L., Snir, S., and Gefen, E. 2015. Scorpion speciation in the Holy Land: Multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among Scorpio subspecies and justifies recognition as phylogenetic, ecological and biological species. Molecular Phylogenetics and Evolution 91: 226–237. PDF




Bryson, R.W., Jr., Prendini, L., Savary, W.E., and Pearman, P.B. 2014. Caves as microrefugia: Pleistocene phylogeography of the troglophilic North American scorpion Pseudouroctonus reddelli. BMC Evolutionary Biology 14(9): 1–16. PDF

Cushing, P.E., Casto, P., Knowlton, E.D., Royer, S., Laudier, D., Gaffin, D.D., Prendini, L., and Brookhart, J.O. 2014. Comparative morphology and functional significance of setae called papillae on the pedipalps of male camel spiders (Arachnida, Solifugae). Annals of the Entomological Society of America 107(2): 510–520. PDF

Foord, S.H., Gelebe, V., and Prendini, L. 2014. Effects of aspect and altitude on scorpion diversity along an environmental gradient in the Soutpansberg, South Africa. Journal of Arid Environments 113: 114–120. PDF

Loria, S.F. and Prendini, L. 2014. Homology of the lateral eyes of Scorpiones: A six-ocellus model. PLoS One 9(12): e112913 (30 pp.). doi:10.1371/journal.pone.0112913. PDF

Miller, A.L., Makowsky, R.A., Formanowicz, D.R., Prendini, L., and Cox, C.L. 2014. Cryptic genetic diversity and complex phylogeography of the boreal North American scorpion, Paruroctonus boreus (Vaejovidae). Molecular Phylogenetics and Evolution 71: 298–307. PDF

Monod, L. and Prendini, L. 2014. Evidence for Eurogondwana: The roles of dispersal, extinction and vicariance in the evolution and biogeography of Indo-Pacific Hormuridae (Scorpiones: Scorpionoidea). Cladistics (early edition: 10.1111/cla.12067, published 5 FEB 2014): 1–41. PDF

Santibáñez-Lopez, C.E., Francke, O.F. and Prendini, L. 2014. Kolotl n. gen. (Scorpiones: Diplocentridae) a new scorpion genus from Mexico. American Museum Novitates 3815: 1–14. PDF

Santibáñez-Lopez, C.E., Francke, O.F. and Prendini, L. 2014. Phylogeny of the North American scorpion genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) based on morphology, nuclear and mitochondrial DNA. Arthropod Systematics and Phylogeny 72(3): 257–279. PDF

Santibáñez-Lopez, C.E., Francke, O.F. and Prendini, L. 2014. Shining a light into the world’s deepest caves: Phylogenetic systematics of the troglobiotic scorpion genus Alacran Francke, 1982 (Typhlochactidae: Alacraninae). Invertebrate Systematics 28: 643–664. PDF

Tahir, H.M., Navidpour, S., and Prendini, L. 2014. First reports of Razianus (Scorpiones: Buthidae) from Iraq and Pakistan, descriptions of two new species, and redescription of Razianus zarudnyi American Museum Novitates 3806: 1–26. PDF

Tahir, H.M. and Prendini, L. 2014. Redescription of Heterometrus latimanus and confirmation of the genus Heterometrus (Scorpiones: Scorpionidae) in Pakistan. American Museum Novitates 3805: 1–23. PDF




Bryson, R.W., Jr., Riddle, B.R., Graham, M.R., Tilston Smith, B., and Prendini, L. 2013. As old as the hills: Montane scorpions in southwestern North America reveal an ancient association between biotic diversification and landscape history. PLOS One 8(1): e52822 (11 pp.). PDF

Bryson, R.W., Jr., Savary, W.E., and Prendini, L. 2013. Biogeography of scorpions in the Pseudouroctonus minimus complex (Vaejovidae) from south-western North America: Implications of ecological specialization for pre-Quaternary diversification. Journal of Biogeography 40(10): 1850–1860. PDF

González-Santillán, E. and Prendini, L. 2013. Redefinition and generic revision of the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with descriptions of six new genera. Bulletin of the American Museum of Natural History 382: 1–71. PDF

Graham, M.R., Jaeger, J.R., Prendini, L., and Riddle, B.R. 2013. Phylogeography of Beck's desert scorpion, Paruroctonus becki, reveals Pliocene diversification in the Eastern California Shear Zone and postglacial expansion in the Great Basin Desert. Molecular Phylogenetics and Evolution 69: 502–513. PDF

Graham, M.R., Jaeger, J.R., Prendini, L., and Riddle, B.R. 2013. Phylogeography of the Arizona hairy scorpion (Hadrurus arizonensis) supports a model of biotic assembly in the Mojave Desert and adds a new Pleistocene refugium. Journal of Biogeography 40(7): 1298–1312. DOI: 10.1111/jbi.12079. PDF

Monod, L., Harvey, M.S., and Prendini, L. 2013. Stenotopic Hormurus Thorell, 1876 scorpions from the monsoon ecosystems of northern Australia, with a discussion on the evolution of burrowing behaviour in Hormuridae Laurie, 1896. Revue suisse de Zoologie 120(2): 281–346. PDF

Murienne, J., Benavides, L.R., Prendini, L., Hormiga, G., and Giribet, G. 2013. Forest refugia in Western and Central Africa as ‘museums’ of Mesozoic biodiversity. Biology Letters 9: 20120932 (8 pp.). PDF

Ochoa, J.A., Rojas-Runjaic, F., Pinto-da-Rocha, R., and Prendini, L. 2013. Systematic revision of the Neotropical genus Chactopsis Kraepelin, 1912 (Scorpiones: Chactidae), with descriptions of two new genera and four new species. Bulletin of the American Museum of Natural History 378: 1–121. PDF

Santibáñez-Lopez, C.E., Francke, O.F. and Prendini, L. 2013. Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. American Museum Novitates 3777: 1–47. PDF




Engelbrecht, I. and Prendini, L. 2012. Cryptic diversity of South African trapdoor spiders: Three new species of Stasimopus Simon, 1892 (Mygalomorphae, Ctenizidae), and redescription of Stasimopus robertsi Hewitt, 1910. American Museum Novitates 3732: 1–42. PDF

Klußman-Fricke, B.-J., Prendini, L., and Wirkner, C.S. 2012. Evolutionary morphology of the hemolymph vascular system in scorpions: A character analysis. Arthropod Structure & Development 41: 545–560. PDF

Mattoni, C.I., Ochoa, J.A., Ojanguren-Affilastro, A.A., and Prendini, L. 2012. Orobothriurus (Scorpiones: Bothriuridae) phylogeny, Andean biogeography, and the relative importance of genitalic and somatic characters. Zoologica Scripta 41(2): 160–176. PDF

Ojanguren-Affilastro, A.A., Mattoni, C.I., Ochoa, J.A., and Prendini, L. 2012. Rumikiru, n. gen. (Scorpiones: Bothriuridae), a new scorpion genus from the Atacama Desert. American Museum Novitates 3731: 1–43. PDF

Prendini, L. 2012. Scorpions (Scorpiones). In: Grzimek’s Animal Life (N. Simmons, ed.). Gale Cengage. https://animals.galegroup.com Last modified: February 10, 2012. PDF

Sissom, W.D., Hughes, G.B., Bryson, R.W., Jr., and Prendini, L. 2012. The vorhiesi group of Vaejovis C.L. Koch, 1836 (Scorpiones: Vaejovidae) in Arizona, with description of a new species from the Hualapai Mountains. American Museum Novitates 3742: 1–19. PDF

Volschenk, E.S., Harvey, M.S. and Prendini, L. 2012. A new species of Urodacus (Scorpiones: Urodacidae) from Western Australia. American Museum Novitates 3748: 1–18. PDF




Engelbrecht, I. and Prendini, L. 2011. An assessment of the taxonomic resolution of southern African trapdoor spiders (Araneae: Ctenizidae; Cyrtaucheniidae; Idiopidae) and implications for their conservation. Biodiversity and Conservation 20: 3101–3116. PDF

Ochoa, J.A., Ojanguren-Affilastro, A.A., Mattoni, C.I., and Prendini, L. 2011. Systematic revision of the Andean scorpion genus Orobothriurus Maury, 1976 (Bothriuridae), with discussion of the altitude record for scorpions. Bulletin of the American Museum of Natural History 359: 1–90. PDF

Ojanguren-Affilastro, A.A., Pizarro-Araya, J., and Prendini, L. 2011. New data on Chilean Urophonius Pocock, 1893 (Scorpiones, Bothriuridae), with description of a new species. American Museum Novitates 3725: 1–44. PDF

Pizarro-Araya, J., Ojanguren-Affilastro, A.A., and Prendini, L. 2011. First report of an arboreal scorpion (Bothriuridae: Urophonius) from the temperate forests of southern Chile. Gayana 75(2): 166–170. PDF

Prendini, L. 2011. Order Scorpiones C.L. Koch, 1850. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 115–117. PDF

Prendini, L. 2011. Order Solifugae Sundevall, 1833. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 118. PDF

Prendini, L. 2011. Order Palpigradi Thorell, 1888. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 121. PDF

Prendini, L. 2011. Order Ricinulei Thorell, 1876. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 122. PDF

Prendini, L. 2011. Order Amblypygi Thorell, 1883. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 154. PDF

Prendini, L. 2011. Order Thelyphonida Latreille, 1804. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 155. PDF

Prendini, L. 2011. Order Schizomida Petrunkevitch, 1945. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 156. PDF




Mattoni, C., Ochoa, J., Ojanguren-Affilastro, A., and Prendini, L. 2010. Towards an all-species phylogeny of the scorpion family Bothriuridae. Cladistics 26: 216–217.

Ochoa, J.A., Botero-Trujillo, R., and Prendini, L. 2010. On the Troglomorphic scorpion Troglotayosicus humiculum (Scorpiones, Troglotayosicidae), with first description of the adults. American Museum Novitates 3691: 1–19. PDF

Ochoa, J.A. and Prendini, L. 2010. The genus Hadruroides Pocock, 1893 (Scorpiones: Iuridae) in Peru: New records and descriptions of six new species. American Museum Novitates 3687: 1–56. PDF

Ojanguren-Affilastro, A.A., Ochoa, J.A., Mattoni, C.I., and Prendini, L. 2010. Systematic revision of the granulatus group of Urophonius Pocock 1893 (Scorpiones, Bothriuridae) with description of a new species from central Chile. American Museum Novitates 3695: 1-40. PDF

Prendini, L. 2010. Order Scorpiones C.L. Koch, 1837. Pp. 321–330 In: Gerlach, J. and Marusik, Y. (Eds.) Arachnida and Myriapoda of the Seychelles Islands. Siri Scientific Press, Manchester, UK. PDF

Prendini, L. and Esposito, L.A. 2010. A reanalysis of Parabuthus (Scorpiones: Buthidae) phylogeny with a description of two new Parabuthus species endemic to the Central Namib gravel plains, Namibia. Zoological Journal of the Linnean Society 159(3):673-710. PDF

Prendini, L., Francke, O.F. and Vignoli, V. 2010. Troglomorphism, trichobothriotaxy and typhlochactid phylogeny (Scorpiones, Chactoidea): More evidence that troglobitism is not an evolutionary dead-end. Cladistics 26: 117–142. PDF




Dippenaar-Schoeman, A.S., Van den Berg, A. and Prendini, L. 2009. Spiders and scorpions (Arachnida: Araneae, Scorpiones) of the Nylsvley Nature Reserve, South Africa. Koedoe 51(1): 1–9. DOI: 10.4102/koedoe.2009.161. PDF

Francke, O.F., Vignoli, V. and Prendini, L. 2009. A new species of Typhlochactas (Scorpiones, Typhlochactinae) from eastern Mexico. American Museum Novitates 3647: 1–11. PDF

Huff, J.C. and Prendini, L. 2009. On the African whip scorpion, Etienneus africanus (Hentschel, 1899) (Thelyphonida: Thelyphonidae), with a redescription based on new material from Guinea-Bissau and Senegal. American Museum Novitates 3658: 1–17. PDF

Prendini, L., Esposito, L.A., Huff, J.C. and Volschenk, E.S. 2009. Redescription of Rhopalurus abudi Armas & Marcanoi Fondeur (Scorpiones: Buthidae), with first description of the male and first record from Hispaniola. Journal of Arachnology 37: 206–244. PDF

Vignoli, V. and Prendini, L. 2009. Systematic revision of the troglomorphic scorpion family Typhlochactidae (Scorpiones: Chactoidea). Bulletin of the American Museum of Natural History 326: 1–94. Cover PDF




Dunlop, J.A., Tetlie, O.E. and Prendini, L. 2008. Reinterpretation of the Silurian scorpion Proscorpius osborni (Whitfield): Integrating data from Palaeozoic and Recent scorpions. Palaeontology 51(2): 303–320. PDF

Francke, O.F. and Prendini, L. 2008. Phylogeny and classification of the giant hairy scorpions, Hadrurus Thorell (Iuridae Thorell): A reappraisal. Systematics and Biodiversity 6(2): 205-223. PDF

Huff, J.C., Viquez, C., and Prendini, L. 2008. Redescription of Mimoscorpius pugnator (Butler, 1872) (Arachnida: Thelyphonida), with first description of the female. American Museum Novitates 3633: 1–9. PDF

Kamenz, C. and Prendini, L. 2008. An atlas of book lung fine structure in the Order Scorpiones (Arachnida). Bulletin of the American Museum of Natural History 316: 1–259. Popular Press Coverage 1 2 PDF

Ochoa, J.A.and Chaparro, J.C. 2008. A new scorpion species of the genus Hadruroides (Scorpiones: Caraboctoninae) from inter Andean valleys of Peru . Revista Peruana de Biología 15(1): 5-10. PDF

Prendini, L. and Bird, T.L. 2008. Scorpions of the Brandberg Massif, Namibia: Species richness inversely correlated with altitude. African Invertebrates 49(2): 77–107. PDF

Vignoli, V. and Prendini, L. 2008. A new species of Akentrobuthus Lamoral, 1976 (Scorpiones: Buthidae) from the Republic of Benin (Western Africa). Journal of Afrotropical Zoology 4: 61-70. PDF

Volschenk, E.S., Mattoni, C.I. and Prendini, L. 2008. Comparative anatomy of the mesosomal organs of scorpions (Chelicerata, Scorpiones), with implications for the phylogeny of the order. Zoological Journal of the Linnean Society 154: 651-675. PDF

Volschenk, E.S. and Prendini, L. 2008. Aops oncodactylus gen. et sp. nov., the first troglobitic urodacid (Chelicerata: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions. Invertebrate Systematics 22: 235-257. PDF




Baldo, L., Prendini, L., Corthals, A. and Werren, J. 2007. Wolbachia are present in southern African scorpions and cluster with supergroup F. Current Microbiology 55: 367–373. PDF

Francke, O.F. 2007. Scorpions (Arachnida, Scorpiones) from Frontera Corozal, in the Lacandona rainforest, Chiapas, México, with the description of a new species of Diplocentrus (Diplocentridae). Revista Mexicana de Biodiversidad 78: 69- 77. PDF

Francke, O.F., Gonzalez-Santillan, E. 2007. A new species belonging to the Vaejovis puntiplapi group (Scorpiones, Vaejovidae) from southern Mexico. Journal of Arachnology 34:586–591. PDF

Harvey, M.S., Volschenk, E.S. 2007. A review of some Australian Chernitidae: Sundochernes, Troglochernes, and a new genus (Pseudoscorpiones). Journal of Arachnology 35:238–277. PDF

Mattoni, C.I. 2007. The genus Bothriurus (Scorpiones, Bothriuridae) in Patagonia. Insect Systematics and Evolution 38(1): 1-22. PDF

Ochoa, J.A. and Ojanguren Affilastro, A. 2007. Systematics and distribution of Brachistosternus (Brachistosternus) ehrenbergii (Gervais, 1841), with the first record of stridulation in this genus Brachistosternus (Scorpiones: Bothriuridae). Studies on Neotropical Fauna and Environment 42:61–69. PDF

Ojanguren-Affilastro, A.A., Agusto, P., Pizarro-Araya, P., Mattoni, C.I. 2007. Two new scorpion species of genus Brachistosternus (Scorpiones: Bothriuridae) from northern Chile. Zootaxa 1623: 55–68. PDF

Ojanguren Affilastro, A.A., Mattoni, C.I. and Prendini, L. 2007. The genus Brachistosternus (Scorpiones: Bothriuridae), in Chile, with descriptions of two new species. American Museum Novitates 3564: 1–44. PDF

Ramírez, M.J., Coddington, J.A., Maddison, W.P., Midford, P., Prendini, L., Miller, J., Griswold, C.E., Hormiga, G., Sierwald, P., Scharff, N., Benjamin, S.P. and Wheeler, W.C. 2007. Linking of digital images to phylogenetic data matrices using a morphological ontology. Systematic Biology 56(2): 283–294. PDF

Russell, K.N., Do, T.M., Huff, J.C., Platnick, N.I. 2007. Introducing SPIDA-Web: Wavelets, Neural Networks and Internet Accessibility in an Image-Based Automated Identification System. pp. 131-152 In: MacLeod, N. (Ed.) Automated Taxon Identification in Systematics. PDF

Wirkner, C.S. and Prendini, L. 2007. Comparative morphology of the hemolymph vascular system in scorpions – a survey using corrosion casting, MicroCT and 3D-reconstruction. Journal of Morphology 268: 401–413. PDF




Francke, O. F., Savary, W.E. 2006. A new troglobitic Pseudouroctonus Stahnke (Scorpiones: Vaejovidae) from northern México Zootaxa 1302: 21–30. PDF

Goloboff, P.A., Mattoni, C.I. , Quinteros, A.S. 2006. Continuous characters analyzed as such. Cladistics 22: 589–601. PDF

Mattoni, C.I., Acosta, L.E. 2006. Systematics and distribution of three Bothriurus species (Scorpiones, Bothriuridae) from central and northern Chile . Studies on Neotropical Fauna and Environment 41(3): 235–250. PDF

Mercurio, R. 2006. New centipede records from Long Island, New York. Journal of the New York Entomological Society 113(3–4): 232–233. PDF

Ojanguren Affilastro, A.A. and Mattoni, C.I. 2006. A new species of Brachistosternus from Chilean central Andes (Scorpiones, Bothriuridae). Studies on Neotropical Fauna & Environment 41: 79-85. PDF

Prendini, L. 2006. New South African flat rock scorpions (Liochelidae: Hadogenes). American Museum Novitates 3502: 1–32. PDF

Prendini, L. "Scorpiones", in AccessScience, McGraw−Hill, https://www.accessscience.com, DOI 10.1036/1097−8542.607800, last modified: April 7, 2006. PDF

Prendini, L., Volschenk, E.S., Maaliki, S., and Gromov, A.V. 2006. A ‘living fossil’ from Central Asia: The morphology of Pseudochactas ovchinnikovi Gromov, 1998 (Scorpiones: Pseudochactidae), with comments on its phylogenetic position. Zoologischer Anzeiger 245: 211–248. PDF

Roig-Juñent, S., Domínguez, M.C., Flores, G.E., and Mattoni, C.I. 2006. Biogeographic history of South American arid lands. A view from its arthropods. Journal of Arid Environments 66: 404–420. PDF




Mattoni, C.I. 2005. Tergal and sexual anomalies in bothriurid scorpions (Scorpiones, Bothriuridae). Journal of Arachnology 33: 622–628. PDF

Mattoni, C.I. and Acosta L.E. 2005. A new species of Bothriurus (Scorpiones, Bothriuridae) from Brazil. Journal of Arachnology 33: 735-744. PDF

Monod, L. and Lourenço, W.R. 2005. Hemiscorpiidae (Scorpiones) from Iran, with descriptions of two new species and notes on biogeography and phylogenetic relationships. Revue Suisse de Zoologie 112(4): 869–941. PDF

Ochoa, J. A. 2005. Brachistosternus ninapo una nueva especie (Scorpiones: Bothriuridae) de los Andes occidentales en el sur del Perú. Revista Peruana de Biología 11(2):139-148. PDF

Ochoa, J.A. 2005. Patrones de distribución de escorpiones de la región andina en el sur peruano. Revista Peruana de Biología 12(1): 49-68. PDF

Prendini, L. 2005. Comment on “Identifying spiders through DNA barcodes”. Canadian Journal of Zoology 83(3): 481–491. PDF

Prendini, L. 2005. New records and observations on the natural history of Lisposoma elegans and Lisposoma josehermana (Scorpiones: Bothriuridae). American Museum Novitates 3487: 1-11. PDF

Prendini, L. 2005. On Hadogenes angolensis Lourenço 1999 syn. n. (Scorpiones, Liochelidae), with a redescription of H. taeniurus (Thorell 1876). Revue Suisse de Zoologie 112(2): 1–28. PDF

Prendini, L. 2005. Scorpion diversity and distribution in southern Africa: Pattern and process. pp. 25–68 In: Huber, B.A., Sinclair, B.J. & Lampe, K.-H. (Eds.) African Biodiversity: Molecules, Organisms, Ecosystems. Proceedings of the 5th International Symposium on Tropical Biology, Museum Alexander Koenig, Bonn. Springer Verlag, New York. PDF

Prendini, L. and Wheeler, W.C. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics 21: 446-494. PDF

Prendini, L., Weygoldt, P. and Wheeler, W.C. 2005. Systematics of the Damon variegatus group of African whip spiders (Chelicerata: Amblypygi): Evidence from behaviour, morphology and DNA. Organisms, Diversity and Evolution 5: 203–236. PDF

Shelley, R.M. and Mercurio, R. 2005. Ectonocryptoides quadrimeropus, a new centipede genus and species from Jalisco, Mexico; proposal of Ectonocryptopinae, analysis of subfamilial relationships, and a key to subfamilies and genera of Scolopocryptopidae (Scolopendromorpha). Zootaxa 1094: 25–40. PDF

Vignoli,V., Salomone, N., Caruso, T., and Bernini, F. 2005. The Euscorpius tergestinus (C.L. Koch, 1837) complex in Italy:Biometrics of sympatric hidden species (Scorpiones: Euscorpiidae) Zoologischer Anzeiger 244: 97–113. PDF

Volschenk, E.S. 2005. A new technique for examining surface morphosculpture of scorpions. Journal of Arachnology 33: 820-825. PDF




Coddington, J.A., Giribet, G., Harvey, M.S., Prendini, L., and Walter, D.E. 2004. Arachnida. pp. 296–318 In: Cracraft, J. and Donoghue, M. (Eds.) Assembling the Tree of Life. Oxford University Press, Oxford. PDF

Druce, D., Hamer, M., Slotow, R. and Prendini, L. 2004. Checklist of millipedes (Diplopoda), centipedes (Chilopoda) and scorpions (Arachnida: Scorpionida) from a savanna ecosystem, Limpopo Province, South Africa. African Invertebrates 45: 315–322. PDF

González Santillán, E. 2004. Diversidad, taxonomía y hábitat de alacranes. pp. 25-35 In: García, A.N. & Ayala R. (Eds.) Artrópodos de Chamela. Institito de Biologia, UNAM, México. PDF

González Santillán, E., Sissom, W.D., and Pérez, T.M. 2004. Description of the male of Vaejovis sprousei Sissom, 1990 (Scorpiones: Vaejovidae). Texas Memorial Museum, Speleological Monographs 6: 9-12. PDF

Goloboff, P.A., Mattoni, C.I. and Quinteros, A.S. 2004. Continuous characters analyzed as such. Cladistics 20: 595. PDF

Isbister, G.K., Volschenk, E.S., and Seymour, J.E. 2004. Scorpion stings in Australia: Five definite stings and a review. Internal Medicine Journal 34: 427–430. PDF

Mattoni, C.I. and Peretti A.V. 2004. The giant and complex genital plug of the asper group of Bothriurus (Scorpiones, Bothriuridae): morphology and comparison with other genital plugs in scorpions. Zoologischer Anzeiger 243(1-2): 75-84. PDF

Monod, L. and Volschenk, E.S. 2004. Liocheles litodactylus (Scorpiones: Liochelidae): An unusual new Liocheles species from the Australian Wet Tropics (Queensland). Memoirs of the Queensland Museum 49(2): 675–690. PDF

Prendini, L. 2004. On the Scorpions of Gabon and neighboring countries, with a reassessment of the synonyms attributed to Babycurus buettneri Karsch and a redescription of Babycurus melanicus Kovaík. California Academy of Sciences Memoir 28: 235–267. PDF

Prendini, L. 2004. Revision of Karasbergia Hewitt (Scorpiones: Buthidae), a monotypic genus endemic to southern Africa. Journal of Afrotropical Zoology 1: 77–93. PDF

Prendini, L. 2004. Systematics of the genus Pseudolychas Kraepelin (Scorpiones: Buthidae). Annals of the Entomological Society of America 97(1): 37-63. PDF

Prendini, L. 2004. The systematics of southern African Parabuthus Pocock (Scorpiones, Buthidae): Revisions to the taxonomy and key to the species. Journal of Arachnology 32(1): 109–186. PDF

Prendini, L., Crowe, T.M., and Wheeler, W.C. 2004. Burrowing scorpion phylogeny and the evolution of paleodrainage systems in southern Africa. Cladistics 20: 93. PDF

Prendini, L., and Wheeler, W.C. 2004. Assembling the Scorpion Tree of Life: Phylogeny of extant scorpions based on molecules, morphology and exemplars. Cladistics 20: 602–603. PDF

Ochoa, J. A. 2004. Filogenia del género Orobothriurus y descripción de un nuevo género de Bothriuridae (Scorpiones). Revista Ibérica de Aracnología 9:43-73. PDF

Sissom, W.D. and González Santillán, E. 2004. A new species and new records for the Vaejovis nitidulus group, with a key to the Mexican species (Scorpiones, Vaejovidae). Texas Memorial Museum, Speleological Monographs 6: 1-8. PDF




Acosta, L.E. and Mattoni, C.I. 2003. Scorpio chilensis Molina, 1782 (currently Bothriurus chilensis: Arachnida, Scorpiones): Proposed suppression of the specific name. Case 3246. Bulletin of Zoological Nomenclature 60(2): 113–116. PDF

Fet, V., Soleglad, M.E., Gantenbein, B., Vignoli, V., Salomone, N., Fet, E.V. and Schembri, P.J. 2003. New molecular and morphological data on the “Euscorpius carpathicus” species complex (Scorpiones: Euscorpiidae) from Italy, Malta, and Greece justify the elevation of E. c. sicanus (C. L. Koch, 1837) to the species level. Revue Suisse de Zoologie 110: 355-379. PDF

Isbister, GK, Volschenk, E.S., Balit, C.R., and Harvey, M.S. 2003. Australian scorpion stings: A prospective study of definite stings. Toxicon 41: 877–883. PDF

Ochoa, J. A. 2003. Nueva especie de Brachistosternus Pocock (Scorpiones: Bothriuridae) del sur del Perú. Revista Peruana de Biología 9(2):55-63. PDF

Ochoa, J. A. and Acosta, L. E. 2003. Una nueva especie de Orobothriurus (Scorpiones: Bothriuridae) del Santuario Nacional Ampay, Apurímac, Perú. Revista Peruana de Entomología 43:1-6. PDF

Prendini, L. 2003. A new genus and species of bothriurid scorpion from the Brandberg Massif, Namibia, with a reanalysis of bothriurid phylogeny and a discussion of the phylogenetic position of Lisposoma Lawrence. Systematic Entomology 28: 1–24. PDF

Prendini, L. 2003. Discovery of the male of Parabuthus muelleri, and implications for the phylogeny of Parabuthus (Scorpiones: Buthidae). American Museum Novitates 3408: 1–24. PDF

Prendini, L. 2003. Revision of the genus Lisposoma Lawrence 1928 (Scorpiones: Bothriuridae). Insect Systematics and Evolution 34: 241–264. PDF

Prendini, L., Crowe, T.M. and Wheeler, W.C. 2003. Systematics and biogeography of the family Scorpionidae (Chelicerata: Scorpiones), with a discussion of phylogenetic methods. Invertebrate Systematics 17(2): 185–259. PDF

Roig-Juñent, S., Flores, G.E. and Mattoni, C.I.. 2003. Consideraciones biogeográficas de la Precordillera (Argentina), basadas en artrópodos epígeos. pp. 275-288, In: Una perspectiva latinoamericana de la Biogeografía, J.J. Morrone and J. Llorente Bousquets (eds.), México D.F., México. PDF

Vignoli, V., Kovařík, F. and Crucitti, P. 2003. Scorpiofauna of Kashan (Esfahan Province, Iran) (Arachnida, Scorpiones). Euscorpius 9: 1–7. PDF




Acosta, L. E. and Ochoa J. A. 2002. Lista de los escorpiones bolivianos (Chelicerata: Scorpiones), con notas sobre su distribución. Revista de la Sociedad Entomológica Argentina 61(3-4):15-23. PDF

Burke, R.L. and Mercurio, R.J. 2002. Food habits of a New York population of Italian wall lizards, Podarcis sicula (Reptilia, Lacertidae). American Midland Naturalist 147: 368–375. PDF

Dyason, K., Brandt, W., Prendini, L., Verdonck, F., Tytgat, J., Du Plessis, J., Müller, G. and Van der Walt, J. 2002. Determination of species-specific components in the venom of Parabuthus scorpions from southern Africa using matrix-assisted laser desorption time-of-flight mass spectrometry. Rapid Communications in Mass Spectrometry 16: 768–773. PDF

Harvey, M.S. and Volschenk, E.S. 2002. A forgotten scorpion: The identity of Buthus flavicruris Rainbow, 1896 with notes on Urodacus manicatus (Thorell). Records of the Western Australian Museum 21: 105–106. PDF

Mattoni, C.I. 2002. Bothriurus picunche sp. nov., a new scorpion from Chile (Bothriuridae). Studies on Neotropical Fauna and Environment 37(2): 169-174. PDF

Mattoni, C.I. 2002. Bothriurus pichicuy sp. nov., nuevo escorpión chileno del grupo vittatus (Scorpiones: Bothriuridae). Iheringia, Ser. Zool. 92(4): 81-87. PDF

Mattoni, C.I. 2002. La verdadera identidad de Bothriurus vittatus (Guérin Méneville, [1838]) (Scorpiones, Bothriuridae). Revue Arachnologique 14(5): 59-72. PDF

Nishiguchi, M.K., Doukakis, P., Egan, M., Goldstein, P.Z., Kizirian, D., Phillips, A., Prendini, L., Rosenbaum, H.C., Torres, E., Wyner, Y., DeSalle, R. & Giribet, G. 2002. DNA isolation procedures. pp. 249-287 In: DeSalle, R., Giribet, G. and Wheeler, W.C. (Eds.) Methods and Tools in Biosciences and Medicine. Techniques in Molecular Evolution and Systematics. Birkhäuser Verlag AG, Basel. PDF

Ochoa, J. A. and Acosta, L. E. 2002. Orobothriurus atiquipa, a new Bothriurid species (Scorpiones) from Lomas in Southern Peru. The Journal of Arachnology 30:98-103. PDF

Ochoa, J. A. and Acosta, L. E. 2002. Two New Andean Species of Brachistosternus Pocock (Scorpiones: Bothriuridae). Euscorpius: Occasional Publications in Scorpiology 2:1-13. PDF

Prendini, L., Hanner, R. and DeSalle, R. 2002. Obtaining, storing and archiving specimens and tissue samples for use in molecular studies. pp. 176-248 In: DeSalle, R., Giribet, G. and Wheeler, W.C. (Eds.) Methods and Tools in Biosciences and Medicine. Techniques in Molecular Evolution and Systematics. Birkhäuser Verlag AG, Basel. PDF1 PDF2 PDF3 PDF4 PDF5




Monod, L. and Lourenço, W.R. 2001. A new species of Broteochactas Pocock, 1890 from Brazilian Amazonia (Scorpiones: Chactidae). pp. 195–202 In: Fet, V. and Selden, P.A. (Eds.) Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks, UK. PDF

Prendini, L. 2001. A review of synonyms and subspecies in the genus Opistophthalmus C.L. Koch (Scorpiones: Scorpionidae). African Entomology 9(1): 17–48. PDF

Prendini, L. 2001. Book reviews: New volumes on the “minor” arachnid orders. Journal of Arachnology 29(3): 234–237. PDF

Prendini, L. 2001. Further additions to the scorpion fauna of Trinidad and Tobago. Journal of Arachnology 29(2): 173–188. PDF

Prendini, L. 2001. Phylogeny of Parabuthus (Scorpiones, Buthidae). Zoologica Scripta 30(1): 13–35. PDF

Prendini, L. 2001. Species or supraspecific taxa as terminals in cladistic analysis? Groundplans versus exemplars revisited. Systematic Biology 50(2): 290–300. PDF

Prendini, L. 2001. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. pp. 113–138 In: Fet, V. and Selden, P.A. (Eds.) Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks, UK. PDF

Prendini, L. 2001. Two new species of Hadogenes (Scorpiones, Ischnuridae) from South Africa, with a redescription of Hadogenes bicolor and a discussion on the phylogenetic position of Hadogenes. Journal of Arachnology 29(2): 146–172. PDF

Volschenk, E.S., Locket, N.A., and Harvey, M.S. 2001. First record of a troglobitic ischnurid scorpion from Australia (Scorpiones: Ischnuridae). pp. 161–170 In: Fet, V. and Selden, P.A. (Eds.) Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks, UK.




Acosta, L. E. and Ochoa, J. A. 2000. Nueva especie de Orobothriurus Maury del Perú (Scorpiones, Bothriuridae). Revue Arachnologique 13(10):135-144. PDF

Lourenço, W.R. and Monod, L. 2000. Description of a new genus and species of scorpion (Bothriuridae) from Brazil. Ekológia (Bratislava) 19(3): 145–152. PDF

Mercurio, R.J. 2000. Notes on the “Trapdoor” spider Idiops Perty, 1833. Journal of the British Tarantula Society 15(4): 124–129. PDF

Prendini, L. 2000. A new species of Parabuthus Pocock (Scorpiones, Buthidae), and new records of Parabuthus capensis (Ehrenberg), from Namibia and South Africa. Cimbebasia 16: 201–214. PDF

Prendini, L. 2000. Chelicerata (Scorpiones). In: Kirk-Spriggs, A.H. and Marais, E. (Eds.) Dâures – Biodiversity of the Brandberg Massif, Namibia. Cimbebasia Memoir 9: 109–120. PDF

Prendini, L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): An exemplar approach. Cladistics 16(1): 1–78. PDF

Volschenk, E.S., Smith, G.T., and Harvey, M.S. 2000. A new species of Urodacus from Western Australia, with additional descriptive notes on Urodacus megamastigus (Scorpiones). Records of the Western Australian Museum 20: 57–67. PDF




Lourenço, W.R. and Monod, L. 1999. A new species of Ananteris from French Guyana (Scorpiones, Buthidae). Revue suisse de Zoologie 106(2): 301–306. PDF

Lourenço, W.R. and Monod, L. 1999. Confirmation de la validité du genre Hormiops Fage, 1933 avec redescription d’Hormiops davidovi Fage, 1933 (Scorpiones, Ischnuridae). Zoosystema 21(2): 337–344. PDF

Prendini, L., Crowe, T.M., Linder, H.P. and Wheeler, W.C. 1999. Taking the sting out of scorpion systematics: An outline of progress in understanding the systematics of Opistophthalmus C.L. Koch. American Arachnology 60: 6–7.

Rosso de Ferradás, B. and Mattoni, C.I.. 1999. Ácaros reófilos (Acari: Hydrachnidia) de las Sierras de Córdoba (Argentina). III. Rev. Soc. Entomol. Argen. 58(3-4): 109-127. PDF




Lourenço, W.R. and Monod, L. 1998. Redescription of Compsobuthus rugosulus (Pocock, 1900) (Scorpiones, Buthidae) based on specimens from Pakistan. Revue suisse de Zoologie 105(4): 789–796. PDF

Prendini, L. and Linder, H.P. 1998. Phylogeny of the South African species of restioid leafhoppers (Cicadellidae, Cephalelini). Entomologica Scandinavica 29: 11–18. PDF




Mattoni, C.I. and Acosta, L.E. 1997. Scorpions of the insular sierras in the Llanos District (Province of La Rioja, Argentina) and their zoogeographical links. Biogeographica 73(2): 67–80. PDF

Newlands, G. and Prendini, L. 1997. Redescription of Hadogenes zumpti (Scorpiones: Ischnuridae), an unusual rock scorpion from the Richtersveld, South Africa. South African Journal of Zoology 32(3): 76–81. PDF

Prendini, L. 1997. Two new host-restricted restioid leafhoppers in the genus Cephalelus (Cicadellidae: Cephalelini), with descriptions of the females of C. brevipilus Davies, C. daviesi Davies and C. rawsonia Davies. African Entomology 5(2): 273–281. PDF

Prendini, L., Crowe, T.M. and Linder, H.P. 1997. Challenges to scorpion cladistics: Effects of sex, segmentation and psammophily. Cladistics 13: 177–178. PDF

Prendini, L., Linder, H.P. and Picker, M.D. 1997. Parallel cladogenesis, coevolution and sequential evolution in insect-plant associations: A test case. Cladistics 13: 178–179. PDF




Prendini, L., Theron. L-J., Van der Merwe, K. and Owen-Smith, N. 1996. Abundance and guild structure of grasshoppers (Orthoptera: Acridoidea) in communally grazed and protected savanna. South African Journal of Zoology 31(3): 120–130. PDF




Seymour, J.E., Volschenk, E.S., and Scott B. 1995. Record of the scorpion Liocheles karschii (Keyserling, 1855) from northeast Queensland (Scorpionida: Ischnuridae). Memoirs of the Queensland Museum 38: 532. PDF

• Home > 
• Publications > 

© 2006 Lorenzo Prendini; Last Revised: May 2020 Email