Visiting Scientists

Alexander V. Gromov
Junior Research Scientist, Laboratory of Entomology, Institute of Zoology, Almaty, Kazakhstan
(AMNH Collections Study Grant, 2007; NSF-BS&I Solifugae Grant, 2007–)
Alexander Gromov











Alexander Gromov, a research scientist at the Laboratory of Entomology, Institute of Zoology, Kazakhstan, earned an M.Sc. in Biology from the Kazakh State University in 1994 and has been studying for a Ph.D. in Entomology at the Institute of Zoology since 1995. His research focuses on the arachnid order Solifugae worldwide. Since 1993, he published 11 papers dedicated to the taxonomy and zoogeography of solifuges, travelled to and studied the solifuge collections of 6 major European and Russian museums, and examined solifuge materials loaned from 23 other American, European and Asian museum collections. He accumulated a large collection of solifuges, including 116 species and about 2000 specimens, obtained through fieldwork and exchanges with foreign colleagues, and personally collected solifuges in several countries in the Palaearctic region, including Kazakhstan, Kyrgyzstan, Turkmenistan, Uzbekistan, and the United Arab Emirates. Since 2007, he has been supported as a trainee on the NSF-funded BS&I Solifugae grant (http://www.solpugid.com). He visited the AMNH Arachnid and Myriapod Collections for 7 weeks (25 September–14 November) in 2007, supported by an AMNH Collections Study Grant and by the NSF-BS&I Solifugae Grant, to sort and identify AMNH solpugid holdings and work with co-PI Prendini prioritizing samples for DNA sequencing as part of the objectives of the BS&I grant.


Warren E. Savary
Field Associate, Department of Entomology,
California Academy of Sciences, San Francisco, U.S.A.
(NSF-REVSYS Vaejovidae Grant, 2004–; NSF-BS&I Solifugae Grant, 2007–)
Warren Savary, a compliance officer at the U.S. Food and Drug Administration and Field Associate of the California Academy of Sciences, earned an A.A. in Biology at the College of San Mateo in 1972, a B.A. in Ecology and Systematic Biology at San Francisco State University in 1974, and an M.A. in Ecology and Systematic Biology at the same institution in 1990. His research interests include the systematics and phylogeny of the arachnid orders Solifugae and Scorpiones, and the biogeography of western North America. Since 2004, he served as research collaborator and webmaster for the NSF-funded REVSYS Vaejovidae grant and, since 2007, as research collaborator and webmaster for the BS&I Solifugae grant. He has visited the AMNH Arachnid and Myriapod Collections on several occasions to sort and identify solifuges and vaejovid scorpions and to attend grant-related meetings. During his most recent visit (13–19 October, 2007), Savary worked with co-PI Prendini and visiting scientist, Alexander V. Gromov, prioritizing samples for sequencing as part of the objectives of the BS&I Solifugae grant.
Warren Savary








Martín J. Ramírez, PhD
Investigador Adjunto, CONICET -
Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires, Argentina
(2004–2007)
Martín J. Ramírez

Martín Ramírez received his Ph.D., on the systematics and phylogenetics of anyphaenid spiders from the University of Buenos Aires, Argentina. He spent two years as a postdoctoral fellow at the AMNH, between 2000 and 2001, funded by a Fessenden Research Fellowship and a fellowship from CONICET, working on the morphology and phylogenetics of dionychan spiders. He is currently a Senior Collaborator in the National Science Foundation-funded AToL: Phylogeny of Spiders project, coordinating the morphological atlas, the images database, and collecting morphological data on several spider families. He is also a participant in the Goblin Spider Planetary Biodiversity Inventory, lead by Norman Platnick, and also funded by NSF. During past years, Ramírez visited the AMNH on several occasions, in connection with the AToL Spider Phylogeny project, and processed spider tissues collected by Prendini and collaborators in the Middle East and West Africa.


W. David Sissom, Ph.D
Professor of Biology, West Texas A&M University, Canyon, Texas
(NSF-REVSYS Vaejovidae Grant: 2005–2007)
Dr W. David Sissom is a Professor of Biology at West Texas A&M University, Canyon, Texas, where he is primarily engaged in teaching general biology, zoology, entomology, parasitology, medical entomology, arachnology, and systematics. After receiving his Bachelor’s degree in Zoology from Texas Tech University, David entered graduate school and completed his Master’s degree at Texas Tech University, Lubbock (1980) and his Ph.D. at Vanderbilt University (1985). Prior to his current position, he held academic posts at Texas Tech University (1985, 1986) and Elon Collage, North Carolina (1986–1992). Sissom has authored 66 research publications, focusing mainly on the arachnid order Scorpiones, but has occasionally published on spiders, harvestmen, amblypygids, schizomids, and palpigrades. He has specialized on the North American scorpion fauna, particularly the family Vaejovidae. He authored or coauthored several chapters in the Biology of Scorpions (1990) by G. A. Polis (Ed.), and was a coauthor on the Catalog of the Scorpions of the World 1758–1997 (2000). He is currently involved as co-Principal Investigator on the NSF-REVSYS Vaejovidae grant with Lorenzo Prendini, Oscar Francke, Warren Savary, and Edmundo González Santillán.
W. David Sissom










James C. Cokendolpher
Research Associate & Interim Collections Manager of Invertebrate Collection,
Museum of Texas Tech University, Lubbock
(2006)
James C. Cokendolpher
















James Cokendolpher received his M.S. from Midwestern State University, Texas, in 1978. His thesis was on the crab-spiders of north-central Texas. Since then he has studied most groups of modern arachnids (except palpigrades, solpugids, and Uropygi), some fossil Opiliones and Acari, and a few groups of insects and other animals and plants; resulting in over 125 scientific, reviewed publications. He visited the AMNH Southwestern Research Station several times in the 1970's and 1980's and borrowed and donated many specimens from/to the AMNH for taxonomic studies. In the Fall 2005, James went on the AMNH expedition to recollect the elusive new species of ammoxenid spider from the desert of the southwestern border of Texas. It was not until 2006 that James visited the AMNH (rather like going to the Mecca for arachnology!). This trip was primarily intended to work with the AMNH collections and Dr. Pierre Paquin and Nadine Dupérré (from Québec) on a group of troglobitic dictynid spiders. During this trip the lesser arachnid collections were also briefly examined to assess the holdings and locate material of immediate interest for revisionary work on Opiliones from various localities around the globe and North American nesticid spiders and pseudoscorpions. The survey of the Opiliones collection revealed it needs attention and James plans to return to organize this collection and identify all at least to family level.


Pierre Paquin and Nadine Dupérré
(2006)
Nadine Dupérré completed a B.Sc at Université de Montréal in 1998. Mainly interested in Linyphiidae taxonomy and systematics, Nadine is also an accomplished scientific illustrator. You can see her work in the Spiders of Québec and more recently in the Spiders of North America. Pierre Paquin completed an M.Sc. on the effect of disturbances on soil fauna, a Ph.D in community ecology on the relationship of age of forest and community structure of beetles and spiders at Université de Montréal. Post-doctoral studies were carried out in San Diego and Portland on molecular systematics and biogeography of eyeless spiders of the genus Cicurina. Research interests involve cave spiders (mainly in North America) assessed with both morphology and molecules. Taxonomic group of interest includes Linyphidae (Oaphantes, Oreonetides, Phanetta, Agyneta), Dictynidae (Cicurina) and Leptonetidae. The visit at the AMNH allowed Pierre to study and redescribe the numerous types of eyeless Cicurina, and gather data on several other unknown cave spiders scattered in several families. Pierre and Nadine crossed North America a dozen times (in their famous car) to collect cave and surface species good for both DNA and morphology. They also visited and worked in several other museums (CAS in San Francisco, MCZ in Cambridge, and CNC Ottawa) and built over the years an important collection of specimens preserved for DNA analysis. Pierre and Nadine hope to return to the AMNH to complete the Cicurina revision and other revisions on Linyphiidae.
Pierre Paquin and Nadine Dupérré














Oscar Francke, Ph.D.
Curator, National Arachnid Collection, Instituto de Biología, Universidad Nacional Autonóma de México, Mexico City
NSF-REVSYS Vaejovidae Grant: 2005, 2006)
Oscar Francke
















Dr. Oscar Francke was born in Mexico City, where he received his primary education; he received his secondary education in Lima, Peru; and earned a B.S. degree in Entomology in 1970, and a Ph.D. degree in Zoology in 1976, both from Arizona State University. Oscar spent four weeks at the AMNH in the summer of 1976, curating the scorpion collection. He served as Assistant and then Associate Professor at Texas Tech University from 1976 to 1986. In 1986, he returned to Mexico, left the academic world, and donated a substantial collection of scorpions to the AMNH. He worked in private industry for 12 years and was in semi-retirement in 2000 when several Mexican students expressed interest in scorpions. He became involved in their thesis projects and re-entered the academic world at UNAM, where he currently directs the arachnological research of four Master’s students. His primary research interests are scorpion systematics and biology (life histories, reproductive behavior), although for several years he worked with fire ants (Solenopsis) in Texas. He has publsihed over 60 papers on scorpions and almost 20 on ants. In 2003, he was invited by Dr. Prendini to collaborate on the REVSYS Vaejovidae project (www.vaejovidae.com) and has participated in several field trips with co-PIs by Prendini and Dr. W. David Sissom and their students, as well as visited the AMNH on three separate occasions in 2005 and 2006. Oscar is currently involved in a variety of projects, including a grant to produce distribution maps for the scorpions of medical importance (Centruroides) in Mexico, and the distribution of scorpion stings (morbidity and mortality) by state and municipality. In 2006, over 280,000 people, mostly children under 4 years old, required medical treatment for scorpion envenomation in Mexico; 41 people died as a consequence.


Peter Weygoldt, Ph.D.
Professor Emeritus, Albert-Ludwigs University, Freiburg
(2005)
Peter Weygoldt studied zoology at the Universities in Kiel and Freiburg, Germany. In 1958 he graduated at Christian Albrechts University, Kiel, with a PhD on the embryology of crustaceans and then continued these studies for two years in a post-doctoral position. In 1960, he joined the Free University of Berlin as Assistant Professor and started to study the embryology and reproductive behavior of pseudoscorpions, receiving his Venia Legendi for this work. From 1965, he spent two years as Research Associate and Visiting Professor in the Duke Marine Laboratory, Beaufort, North Carolina, and the Friday Harbor Laboratories, University of Washington, studying crustacean embryology. During a short visit to the Florida Keys he found his first whip spider (Amblypygi), and the study of these arachnids became his main interest from then on. Since 1967, Peter served as Professor at Albert-Ludwigs University, Freiburg. There he continued to study embryology, and later reproductive biology and the systematics of Amblypygi. During several travels to Brazil and other Neotropical countries he became fascinated by Neotropical frogs and, for about 20 years, studied their reproductive biology. Since 1995, Peter retired but continues his studies on the Amblypygi. He has published over 130 papers, two books (on the biology of pseudoscorpions and whip spiders) and several contributions to text-books and compendia (e.g. Traité de Zoologie, Lexikon der Biologie). He and his wife, Sylvia, visited the AMNH in the Fall, 2005, to examine the amblypygid holdings in the collections.
Peter Weygoldt
















Mark Harvey, Ph.D.
Senior Curator, Western Australian Museum, Perth
(2006)
Mark Harvey












Mark Harvey, one of Australia’s leading arachnologists, has a broad research focus and has delved into many different arachnid orders including pseudoscorpions, schizomids, spiders and scorpions. He is also an inveterate cataloguer and has produced two catalogues, the first on pseudoscorpions (1991) and the second on the smaller arachnid orders (2003). Other published volumes include Worms to Wasps: an Illustrated Guide to Australia's Terrestrial Invertebrates (co-authored with A.L. Yen), and the CD-ROM Spiders of Australia: interactive identification to subfamily (co-authored with Robert Raven and Barbara Baehr). He originally studied Zoology at Monash University (Melbourne) and graduated in 1983 with a Ph.D. on pseudoscorpions. After two post-doctoral positions in Canberra and Melbourne, he transferred to the Western Australian Museum in 1989 where he is now Senior Curator. Mark has published over 130 research papers and runs an active laboratory with post-docs, students and technicians. During his visit to the AMNH in October 2006, Mark studied the AMNH pseudoscorpion collection and identified some of the unidentified slide material. Mark enjoys foraging in the bush searching for creepy-crawlies, especially with his wife and two daughters, and playing basketball on Wednesday evenings.


O. Erik Tetlie, Ph.D.
Postdoctoral Associate, Yale University
(2006)
Erik Tetlie received his M.S. equivalent from the University of Oslo, Norway, in 2000, for a study on Norwegian eurypterids (sea scorpions). He later received his Ph.D. at the University of Bristol, UK, for a thesis entitled “Eurypterid phylogeny with remarks on the origin of arachnids”. He is currently working on the most comprehensive collection of fossil fauna ever assembled from the eurypterid beds of eastern North America. These include some of the oldest known fossil scorpions, which brought him into contact with Lorenzo Prendini. Tetlie is also very interested in chelicerate phylogeny, especially the respective positions of Eurypterida and Scorpiones, and the original habitat (terrestrial or marginal marine) of the earliest scorpions. While at the AMNH in May 2006, some problematic fossils from the eurypterid beds were tentatively identified as centipedes by Randy Mercurio. These and the scorpions might suggest the presence of a well-functioning terrestrial habitat in the Upper Silurian.
Erik Tetlie







Sergei I. Golovatch, Ph.D.
Senior Scientist, Institute for Problems of Ecology and Evolution
Russian Academy of Sciences, Moscow
(2002, 2006)
Sergei Golovatch
Sergei Golovatch, a myriapodologist at the Russian Academy of Sciences, Moscow, has visited the AMNH Division of Invertebrate Zoology on several occasions to study the millipede (diplopod) collection, most recently for a week in January 2006. During his latest visit, Golovatch worked with Randy Mercurio, examining identified and unidentified bulk collections of Diplopoda to sort out certain subgroups for a few possible future research projects. However, further sorting-out efforts are still highly topical, because the AMNH holdings are extremely abundant, diverse and interesting on a global scale. Golovatch plans to return again to continue his work.


Rowland Shelley, Ph.D.
Curator of Terrestrial Invertebrates, North Carolina State Museum of Natural Sciences, Raleigh
(2002, 2005, 2007)
Rowland Shelley, a myriapodologist at the North Carolina State Museum of Natural Sciences, has visited the AMNH Division of Invertebrate Zoology, and formerly the Department of Entomology, more than 6 times over the past 30 years. He has also visited the AMNH Southwest Research Station in Portal, Arizona. Shelley’s research focuses on the North American millipede (Diplopoda) fauna and the chilopod order Scolopendromorpha, and the AMNH collection of these arthropods is among the 9 largest on the continent. To date, Shelley has reported AMNH specimens in 52 publications, most of which lay the basic taxonomic foundation for higher level studies on these taxa. Dr. Shelley's most recent visit, during September 2005, resulted in the unexpected discovery of a new Mexican scolopendromorph species, genus, and subfamily among unsorted milliped samples, and a collaborative publication with Randy Mercurio. Further collaboration with Mercurio is underway.
Rowland Shelley




Paula Cushing, Ph.D.
Curator of Invertebrate Zoology, Denver Museum of Nature and Science
(Collections Study Grant, 2005)
Paula Cushing



Paula Cushing, an arachnologist at the Denver Museum of Nature and Science, visited the Division of Invertebrate Zoology, AMNH, for a month from January–February 2005, supported by a Collections Study Grant. Cushing examined the solifuge holdings in the collection to determine morphological characters that may be useful for higher-level phylogenetic reconstruction and sequenced DNA from the solifuge tissue samples acquired by Lorenzo Prendini during recent fieldwork in southern Africa, South America and central Asia. These data form the foundations of collaborative research on solifuges to be conducted in the future.